WOLFE, ANDREA D.*, CHRISTOPHER P. RANDLE, and NIDIA ARGUEDAS. Department of Evolution, Ecology, and Organismal Biology, The Ohio State University, 1735 Neil Avenue, Columbus, OH 43210. - Assessing species boundaries in Hyobanche L. (Orobanchaceae).
Hyobanche L. (Orobanchaceae) is a holoparasitic genus of ca.
eight species endemic to southern Africa from the Cape Floristic
Province northward to Namibia. There is a considerable amount of
overlap in morphological characters among species resembling the most
widely distributed species, H. sanguinea. There is also a high
degree of similarity between H. glabrata and H. rubra,
which differ primarily in the degree of exsertion of reproductive
organs from the mouth of the corolla during anthesis. We investigated
the species boundaries of Hyobanche using nucleotide sequence
data from ITS and rbcL, ISSR markers, and morphological data.
Our emphasis was on new collections from the 2001 field season, which
included intensive population sampling in the Little and Nama Karoo,
populations of H. sanguinea from coastal sand dune and
limestone outcrop systems, and what appears to be a hybrid population
between H. sanguinea and H. atropurpurea. A phylogeny
reconstruction using ITS data reveals that H. rubra and H.
glabrata from the Little Karoo form a distinct clade within a
larger clade of populations from H. rubra and H.
glabrata. Additionally, there appear to be few molecular and
morphological characters separating H. rubra and H.
glabrata. Molecular and morphological data also reveal unique
characters for a taxon initially described as H. sanguinea from
the Agulhas Plain.
Key words: Hyobanche, ISSR, Orobanchaceae, parasitic plant, phylogeny, South Africa