The woody Sonchus alliance (Asteraceae: Lactuceae) in the Macaronesian islands is comprised of 19 species of woody Sonchus and five closely related genera and has been regarded as an outstanding example of adaptive radiation among angiosperms. Previous phylogenetic study based on ITS sequences of nrDNA demonstrated the monophyly of the entire alliance and suggested several phylogenetic relationships among the species within the alliance. Further, the ITS phylogeny suspected the placement of several Sonchus species in the Taeckholmia clade due to ancient hybridization and subsequent fixation of the Taeckholmia type ITS sequences. The cpDNA nocoding regions, trnL intron, trnT-L intergenic spacer, and trnL-F intergenic spacer, were sequenced to independently explore relationships within the alliance and also to understand the role of hybridization during the adaptive radiation of the alliance. The cpDNA phylogeny shows low resolution compared to the ITS phylogeny and reveals many different relationships within the alliance. This discordance could be the result of ancient hybridization and subsequent chloroplast capture and further suggests that hybridization appears to have been much more pervasive in the Sonchus alliance than previously suspected.

Key words: adaptive radiation, trnT-L-F, woody Sonchus alliance. Macaronesia