MILLER, JOE. Department of Biological Sciences, The University of Iowa, Iowa City, Iowa 52242. - Apomictic and polyploid evolution in the Australian arid-zone mulga species complex (Acacia: Fabaceae).
Acacia aneura and its close relatives form a highly variable species
complex commonly known as mulga. This tree species complex is the
dominant vegetation in arid and semi-arid Australia and covers 20% of
the continent. Populations contain a high degree of variability in
leaf, habit and fruit characters. Additionally seperate populations
may contain plants with similar morphological features. Field,
greenhouse and laboratory work have been used to examine genetic
causes of the variation in an effort to build a taxonomic treatment
for the group. The complex contains multiple ploidy levels, including
triploids, tetraploids and pentaploids, and polyembryony is a common
feature in all these polyploids. While hybridization has been
suspected in the complex, microsatellite markers could not find direct
evidence of hybridization in the small sampling used to date.
Microsatellite data identified fixed heterozygozity in populations
with some genetic differences among morphotypes. Progeny arrays of 24
morphotypes indicated that over 95% of the plants have the same
genotype as the mother plant. This accumulating evidence indicates
that the complex is reproducing through facultative apomixis.
Key words: Acacia, apomixis, neoteny, polyembryony, polyploidy, species complex