BATES, PAUL L.1*, LINDA E. WATSON2, and JAMES R. ESTES1. 1School of Biological Sciences, University of Nebraska-Lincoln, Lincoln, Nebraska, 68588-0118 USA; 2Department of Botany, Miami University, Oxford, Ohio, 45056 USA. - Molecular systematics of the Artemisia ludoviciana complex (Asteraceae : Tribe Anthemideae) based on the ITS and ETS regions of rDNA.
Artemisia ludoviciana and its allies comprise a complex of 10
species including 7 subspecies. These are endemic to western North
America west of the Mississippi River from the Aleutian Islands to
Oaxaca Mexico, with greatest diversity in the Pacific Northwest. The
complex is highly diverse morphologically and the various forms
intergrade perhaps as a result of polyploidy, hybridization and/or
recent divergence. Overlap of species boundaries has led to troubled
taxonomy and made the production of keys with species decisive
couplets nearly impossible. We constructed phylogenies using internal
transcribed spacers 1 and 2 (ITS) and 3' end external transcribed
spacer (ETS) regions of nuclear ribosomal DNA (rDNA) to 1) determine
the phylogenetic placement of the complex within the genus
Artemisia, 2) determine the relationship of the native North
American taxa and their putative Eurasian relatives, and 3) examine
relationships amongst the nominal forms. Our analysis resulted in the
North American taxa forming a highly supported clade within series
Artemisia (sensu lato) and show a sister relationship to the Eurasian
taxa sampled including the North American species Artemisia
palmeri. We found a low amount of phylogenetically informative
sites amongst the ITS sequences of the complex and attribute this to a
recent origin of the group.
Key words: Artemisia ludoviciana, Asteraceae, ETS, ITS, phylogeny