Satyria, a genus of neotropical blueberries, is a member of the Andean clade of tribe Vaccinieae (Ericaceae) and is distinguished from other neotropical Vaccinieae by alternately unequal stamens composed of connate filaments of equal length and unequal, rigid anthers with divergent tubules. The genus occurs from southern Mexico, south to northern Bolivia and east to French Guiana and is one of only a few species of neotropical Vaccinieae found in both montane and lowland habitats. Satyria may be critical for understanding the origin and biogeography of the blueberry tribe in South and Central America. Although Satyria has been described as a "distinctive genus" of neotropical blueberries, preliminary molecular phylogenetic analyses in tribe Vaccinieae (matK and nrITS) suggest that Satyria is not monophyletic. The goal of this study was to use molecular data from the chloroplast genes matK, ndhF and rps4, and nrITS for approximately 50 species of blueberries from the Andean clade, (including 12 species of Satyria) and three outgroups to assess the monophyly of Satyria and to determine its phylogenetic position in the Andean Clade. The data were analyzed, individually and in combination, using parsimony. Results show that Satyria is not monophyletic but rather forms two well-supported clades: northern and southern clades that correspond to at least two of the regions of high diversity and endemism identified in neotropical Ericaceae. In the northern clade, species of Satyria from Central America and northern South America (including the widely distributed S. panurensis) form a strongly-supported monophyletic group that is sister to a clade that includes Cavendishia and one species of Thibaudia. In the southern clade three species of Satyria from southern Peru and northern Bolivia are paraphyletic with two species of Thibaudia. Results suggest that the southern species of Satyria have been misidentified and should be transferred from Satyria to Thibaudia.

Key words: biogeography, Ericaceae, neotropical blueberries, Satyria, Vaccinieae