The Mimulus aurantiacus complex, comprising endemic shrubs in Southern California, shows a high degree of phenotypic plasticity. It is currently thought that this high degree of plasticity is due to natural hybridization at the subspecies level. However, our studies of Mimulus aurantiacus plants over the last six years suggest that individuals of ssp. australis producing large, yellow, insect-pollinated flowers can undergo a dramatic phenotypic transition to produce small, red, bird-pollinated flowers. The entire transition takes several years until completion. To our knowledge this is the first case demonstrating morphological and biochemical (pigment) alterations over a considerable time span, which finally lead to a new, discrete and stable form. The transitional zone (what is «classically» termed the hybrid zone) migrates towards the East in the most southern part of California. Intrinsic and extrinsic factors are suggested as forcing this rapid transition process resulting in phenotypic evolution. (Mimulus aurantiacus transition http://217.162.182.115 )

Key words: hybrid zone, Mimulus aurantiacus, phenotypic evolution, phenotypic plasticity, pigment, transition zone