STAUFFER, FRED W.* and PETER K. ENDRESS. Institute of Systematic Botany, University of Zurich, Zollikerstrasse 107, CH-8008 Zurich, Switzerland. - Comparative morphology of the gynoecium in the neotropical tribe Geonomeae (Arecaceae: Arecoideae).
The Geonomeae are a neotropical palm tribe with six genera,
Pholidostachys, Welfia, Calyptronoma,
Calyptrogyne, Asterogyne and Geonoma, and up to
70 species. The morphological structure of the gynoecium in Geonomeae
has been in general neglected. Todate, detailed studies on the
reproductive structures have incorporated only two genera and less
than 6% of the species within the tribe. A tri- or uniovulate
gynoecium with slender, elongate styles has been traditionally
considered as typical for the group. We have studied the flower
morphology and anatomy of 15 species representing all genera of
Geonomeae. The gynoecium of Geonoma shows three free carpels in
early stages of development but later becomes pseudomonomerous. The
other five genera show a more equally developed tricarpellate
gynoecium, the two adaxial carpels being larger than the abaxial
carpel. A central protrusion between the three carpels is present in
some species of Pholidostachys, Calyptrogyne and
Geonoma. The ovules in the tribe are attached at the base of
the ovarial cavity or are pendent. They are more or less anatropous.
In all Geonomeae a septal nectary is formed by the incomplete union of
the flanks of the carpels at the base and mid-level of the gynoecium.
Nectar is secreted from an epithelium, and non-secretory ducts are
located at different levels of the ovary. Styles seem to emerge from
the "base" of the ovary, because of the bulging of one of
the carpels in pseudomonomerous gynoecia, but from the top in gynoecia
with all three carpels equally developed. Protuberances in the dorsal
walls of the styles were observed in Pholidostachys,
Calyptrogyne and Asterogyne. The stigmas expose
unicellular papillae at anthesis, in Welfia also multicellular
papillae. Pollen tube transmitting tracts and a compitum are present
in the ventral slits of the postgenitally united styles during
anthesis.
Key words: Arecaceae, Geonomeae, gynoecium, morphology