BROCK, MARCUS T. Division of Biological Sciences, University of Missouri-Columbia, 216 Tucker Hall, Columbia, MO 65211. - The threat of genetic assimilation to the integrity of a native plant species, Taraxacum ceratophorum (Asteraceae).
Invasive exotic species may pose threats to native congeners through
the process of genetic assimilation, where asymmetrical hybridization
and subsequent hybrid backcrossing results in the loss of pure native
populations. I am studying the threats of genetic assimilation by the
exotic Taraxacum officinale into populations of the native
alpine dandelion, Taraxacum ceratophorum. Using expected
mendelian ratios based on allozyme variation, I demonstrated that,
unlike the apomictic exotic T. officinale, populations of T.
ceratophorum on Pennsylvania Mountain (Park County, CO) reproduce
sexually. Initial surveys determined that ecological conditions
required for hybridization between T. ceratophorum and T.
officinale occur for sympatric populations in the Colorado Rocky
Mountains. On Pennsylvania Mountain, native and exotic dandelions
exhibit broad phenological overlap in flowering during 1999 and 2000.
In mixed arrays, insect visitors demonstrated no preference for native
inflorescences over exotic inflorescences. Furthermore, insects move
pollen analogues from exotics to natives and between natives at the
same frequency. Hand-pollination of T. ceratophorum with T.
officinale pollen produced hybrid seeds. To test for reproductive
barriers to the formation of hybrids, I performed three hand
pollination treatments on native inflorescences—a control of native
pollen, native and exotic pollen applied at the same time, and exotic
pollen preceding native pollen by 15 minutes. Hybrids were
consistently produced at low frequencies only when exotic pollen
preceded native pollen by at least 15 minutes. Despite the exotic
pollen disadvantage, other mechanisms may promote the establishment of
hybrids formed in natural populations. Germination rates for parental
and hybrid seeds sown into natural habitats on Pennsylvania Mountain
varied significantly with genotype. Hybrids had a 66% selective
advantage over the native species at this initial life history stage,
which may eventually facilitate backcrossing with the native species.
Key words: genetic assimilation, hybridization, Taraxacum ceratophorum (Asteraceae), Taraxacum officinale (Asteraceae)