BURKE, JOHN M.* and LOREN H. RIESEBERG. Indiana University, Department of Biology, Bloomington, IN 47405. - Genetic analysis of sunflower domestication.
Quantitative trait loci (QTLs) controlling phenotypic differences
between cultivated sunflower and its wild progenitor were investigated
in an F3 mapping population. Composite interval mapping revealed the
presence of seventy-eight QTLs affecting the eighteen quantitative
traits of interest, with two to ten QTLs per trait. Each QTL explained
3.0–68.0% of the phenotypic variance, although only four
(corresponding to three of eighteen traits) had effects > 25%.
Overall, fifty-one of the seventy-eight QTLs produced phenotypic
effects in the expected direction, and for thirteen of eighteen traits
the majority of QTLs had the expected effect. Despite being
distributed across fifteen of the seventeen linkage groups, there was
a substantial amount of clustering among QTLs controlling different
traits. In several cases, regions influencing multiple traits harbored
QTLs with antagonistic effects, producing a cultivar-like phenotype
for some traits and a wild-like phenotype for others. Based on the
directionality of QTLs, strong directional selection for increased
achene size appears to have played a central role in sunflower
domestication. None of the other traits show similar evidence of
selection. The occurrence of numerous wild alleles with cultivar-like
effects, combined with the lack of major QTLs, suggests that sunflower
was readily domesticated.
Key words: domestication, Helianthus, QTL mapping, selection, sunflower