Several fertile pinnules with abaxially borne sporangia have been discovered in calcareous marine nodules from the Late Eocene Appian Way locality on the East coast of Vancouver Island, British Columbia, Canada. Each fertile segment is rolled toward the abaxial surface, forming a roughly spheroidal structure. Sporangia are abaxial and enclosed by the lamina. The specimens also display false indusia that each cover several sporangia. The receptacle is thick and branched, bearing broadly attached sporangia and numerous paraphyses. As is diagnostic for schizaeaceous ferns, sporangia of the fossil are flask-shaped with a short stalk, and have a uniseriate, transverse annulus at the apex. Spores are tetrahedral and trilete, with a psilate surface. The false indusium and unornamented trilete spores suggest affinities with Lygodium, but the grouped sporangia and rolled pinnules of the fossil comprise a unique combination of characters.

Key words: anatomy, Eocene, fern, fossil, Schizaeaceae