GROTE, PAUL J. School of Biology, Institute of Science, Suranaree University of Technology, Nakhon Ratchasima 30000, Thailand. - Tertiary floras of Northern Thailand: additional conifer diversity from charcoalified wood.
Charcoalified fragments of wood scattered in what appear to be
compressed ashes from a fire have been found in Lampang Basin, Lampang
district, Northern Thailand, an inland terrestrial basin of Tertiary
age, possibly Miocene. Approximately four species of conifers are
represented by the charcoalified wood fragments; annual rings are
present in all the wood. One fragment appears to be compression wood,
with tracheids showing extensive splitting of the secondary cell
walls. A traumatic resin canal is also present. A few axial parenchyma
cells were observed, and the rays are uniseriate and biseriate,
consisting of parenchyma. Axial tracheids bear a single row of
bordered pits with subcircular apertures. The crossfields contain one
or two half-bordered pit pairs with oblique elliptic apertures on the
tracheid side. This wood may have affinity with Cupressaceae s.l.
(including Taxodiaceae). A second wood type consists of
thick-walled tracheids, some axial parenchyma, and uniseriate rays
with a height of 36 or more cells. Bordered pits, occurring in one,
two, or possibly more rows along the tracheids, are characterized by
deep pit canals resembling compressed funnels and shallower pit
cavities. One or two half-bordered pit pairs were observed in the
crossfields. This wood type appears to show affinity with
Podocarpaceae, or possibly Cupressaceae. A third wood type shows
affinity to Podocarpaceae, and a possible fourth wood type is similar
to the second type but has shallower, less compressed, pit canals. The
diversity of conifer wood, along with that of other conifer fossils,
twigs, leaves, seed cones, and seeds, from the Tertiary basins of
Northern Thailand, provides evidence that the conifers in the Recent
of Thailand represent a relict of a once larger number of conifers.
Key words: charcoalified wood, conifer, Cupressaceae, Podocarpaceae, Thailand