KRIEGER, JONATHAN D.1*, ROBERT P. GURALNICK1, and KIRK R. JOHNSON2. 1Department of E.P.O. Biology, University of Colorado-Boulder, Boulder, CO 80309; 2Denver Museum of Natural History and Earth Science, Denver, CO 80205. - Morphometric analysis of extant and fossil leaf assemblages.
Many workers have been interested in creating a quantitative means to
determine ecosystem type from aspects of leaf form. We performed
eigenshape analysis of leaf outlines for two extant and two fossil
assemblages. For both the extant and fossil flora, one assemblage
represented leaves from tropical areas and one from temperate. The
data for the extant assemblages suggests that temperate assemblages
occupy a larger region of morphospace. This holds true for the first
twenty eigenshape axes examined. Eigenshape axis 2 appears to
represent the length to width ratio; the tropical leaves are clustered
from an average leaf shape to one with a high length to width ratio,
while the temperate leaves are scattered across the entire axis.
Eigenshape axis 3 appears to show variation due to an elongate tip or
petiole; on this axis, the tropical leaves are clustered near the
average leaf shape. This distribution may reflect the very long
petioles on some temperate leaves; however, it seems to contradict the
expectation that tropical leaves will display longer drip tips. The
combined data for all assemblages agrees with the taxonomic
determination of ecosystem type for the fossil assemblages. The
temperate fossil flora occupies a greater region of morphospace, while
the fossil rainforest flora falls within the morphospace of the extant
tropical flora. The implications of our findings in understanding
fossil assemblages will be discussed, and the broader questions of
paleoclimate determination will be considered.
Key words: climate, ecosystem, fossil assemblage, leaf form, morphometrics