GRAHAM, JAMES M.*, ANGELA D. KENT, GEORGE H. LAUSTER, ANTHONY C. YANNARELL, LINDA E. GRAHAM, TIMOTHY K. KRATZ, and ERIC W. TRIPLETT. Center for Limnology, Limnology Laboratory, 680 N. Park St., University of Wisconsin, Madison, WI 53706-1492. - Species diversity of microbial communities in a northern temperate humic lake.
Species richness and species diversity of bacterioplankton,
phytoplankton and protoplankton were examined in Crystal Bog, a humic
lake in northern Wisconsin, as part of an NSF sponsored Microbial
Observatory Program. Crystal Bog is part of the North Temperate Lakes
Long-Term Ecological Research site. Automated ribosomal intergenic
spacer analysis (ARISA) was used to assess bacterial diversity.
Phytoplankton and protoplankton were enumerated in a settling chamber
with an inverted microscope. Four distinct phases in the ice-free
season of the bog can be recognized based on microbial populations.
The first phase extends from ice-out until about the first week of
April. During this phase three species of dinoflagellates come to
dominate the phytoplankton, with Glenodinium quadridens
representing 60% of total phytoplankton biovolume. By mid-April the
spring dinoflagellate blooms collapse. The second phase is a period of
instability as a series of species dominates the phytoplankton. In
mid-July dinoflagellates again dominate the third phase, and
Peridinium limbatum alone represents 90% of the bog
phytoplankton. The fourth phase is a stable period as P.
limbatum slowly declines in late summer and fall. A strong
correlation was found between bacterial and phytoplankton diversity
(Shannon-Weaver Index) in this bog, suggesting the two microbial
communities are linked. The protoplankton, however, were relatively
uniform throughout the ice-free period. The ice-covered period may be
treated as a fifth phase. When snow accumulates over the ice-covered
bog, light penetration is blocked and photosynthesis shut off.
Heterotrophic protozoa assume dominance in the plankton.
Key words: ARISA, bacteria, bog, microbial diversity, phytoplankton, protozoa