RABELER, RICHARD K.1* and MOLLY NEPOKROEFF2. 1University of Michigan Herbarium, 3600 Varsity Drive Suite 112, Ann Arbor, MI 48108-2287; 2Department of Biology, University of South Dakota, 414 East Clark St., Vermillion, SD 57069-2390. - Systematics and evolution of the Caryophyllaceae: a model family for integrating taxonomic, phylogenetic, and population studies.
The family Caryophyllaceae includes about 85 genera and some 3000
species. Despite much interest in the relationship of the family
within the order Caryophyllales and the core eudicots, only a few
phylogenetic investigations of relationships within the
Caryophyllaceae have been initiated. The current subfamily
classification is based on morphological characters which may be
highly "labile" or homoplastic, such as type of locule
dehiscence. Recently, there has been a surge of interest in the family
at several levels. Researchers using both morphology and molecular
data are now looking at both delimitation of large genera and
relationships within the family. A number of species in the family
have historically been, and continue to be, used as model systems for
investigating the genetic basis of a variety of traits and
evolutionary processes. This colloquium brings together researchers
from North America and Europe to report on their recent findings,
foster collaboration, and discuss questions about classification and
evolution within the family Caryophyllaceae. Are there three, four or
five subfamilies and how can they be recognized? Can large,
polyphyletic genera be split logically based on morphological and
molecular criteria? Are there additional areas of the family sorely in
need of further work? Can phylogenetic and population studies be
better integrated to reflect the explosion in data on both sides? The
first talks will provide an overview of the historic treatment of the
Caryophyllaceae as well as the recent findings about relationships
within the family. The next presentations will focus on evolution and
relationships within specific genera of Caryophyllaceae. The final
talks will focus on studies using members of the Caryophyllaceae as
model systems to study evolutionary phenomena, including studies of
breeding system evolution, gene expression and the molecular genetic
basis for phenotypic traits.
Key words: Caryophyllaceae, classification, evolution, phylogeny, population studies