With the advent of molecular data - and particularly the use of molecular clocks when warranted - biogeographers are re-examining the roles of Gondwana break-up, immigration from the Laurasian tropics, and transoceanic dispersal in shaping similarities among the world's tropical floras. As increasing numbers of continental disjunctions are analyzed, it has also become possible to ask whether there are regular patterns in the direction(s) and timing of transoceanic dispersal events. Wind and ocean surface currents (as well as tectonics) all affect dispersal and are not randomly distributed in space and time; they should thus leave an evolutionary trace provided that they have been stable long enough to override lineage-specific differences in dispersal and establishment capability. This symposium will assemble a diverse set of researchers with phylogenetic data on disjunctly distributed tropical lineages. The mix emphasizes continental disjunctions and lineages that collectively display a variety of dispersal mechanisms, and each contributor has been asked to address three questions: (1) How much information about the timing and nature of the events causing disjunctions can be obtained from molecular data? (2) What appear to be the relative roles of dispersal and vicariance, and how do these relate to aspects of earth history? And (3) How can we crosscheck our inferences? This symposium honors Robert Thorne for his outstanding contributions to biogeography and especially the analysis of tropical plant disjunctions

Key words: angiosperms, boreotopics, Gondwana, Laurasia, long distance dispersal, molecular clocks